Journal
GEODIVERSITAS
Volume 33, Issue 4, Pages 625-635Publisher
MUSEUM NATL HISTOIRE NATURELLE
DOI: 10.5252/g2011n4a4
Keywords
Insecta; Dictyoptera; Pennsylvanian; Late Carboniferous; wing venation pattern; arculus; vannus
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Funding
- RBINS
- European Community
- German Science Foundation [DFG SCHN 408/12]
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The holotype of Phyloblatta gaudryi (Agnus, 1903), provided from the famous deposit of Commentry (Allier, France), exhibits a pair of forewings and a hind wing in connection, well preserved. Forewings show a network of veinlets in the area between M and CuA, and evidence of a fusion of some branches of M with CuA. It is concluded that the m-cua veinlets belong to M and that a reticulated fusion between M and CuA occurs. This fusion is considered as a homologue of the dictyopteran forewing arculus, but as an intermediate polymorphic state between occurrence of a single discrete arculus and no arculus. The preserved hind wing is devoid of an m-cua arculus, has a multi-branched AA vein on the remigium, and has a vannus of limited extent. The new data on P gaudryi might allow the phylogenetic position of Pennsylvanian stem-dictyopterans with respect to crown-dictyopterans to be better assessed.
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