Anisodon(Perissodactyla, Chalicotheriinae) from the middle Miocene locality Gracanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina)

The fossil mammal fauna of the Gracanica coal mine in central Bosnia and Herzegovina includes dental remains ofAnisodon(Perissodactyla, Chalicotheriidae, Chalicotheriinae). These teeth most closely resemble those ofAnisodon grandefrom Sansan (France, MN6) and Neudorf an der March (Slovakia, MN6), and are tentatively referred to that species.Anisodonis first known in Europe from the Loire Basin (France), which has been dated as MN5. Given that the Gracanica fauna is considered to be MN5 or earliest MN6 (Langhian), the Gracanica chalicothere represents one of the earlier known finds of both the Chalicotheriinae andAnisodonin Europe. The first known members of the Chalicotheriinae are from the earliest Miocene of Asia. By the middle Miocene,Anisodonhad appeared and is known from Asia and Europe, to which it had recently immigrated.