4.4 Article

The presence of the recurrent veinlet in the Middle Jurassic Nymphidae (Neuroptera): a unique character condition in Myrmeleontoidea

Journal

ZOOKEYS
Volume -, Issue 325, Pages 1-20

Publisher

PENSOFT PUBL
DOI: 10.3897/zookeys.325.5453

Keywords

Neuroptera; Nymphidae; recurrent veinlet; subcosta anterior; Daohugou; Middle Jurassic

Categories

Funding

  1. National Basic Research Program of China (973 Program) [2012CB821906]
  2. National Science Foundation of China [31230065, 31272352, 41272006]
  3. Project of Great Wall Scholar
  4. KEY project of Beijing Municipal Commission of Education Project [KZ201310028033]
  5. China Geological Survey [1212011120115]

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A well-developed recurrent veinlet is found in the forewing of two species of Nymphidae from the Middle Jurassic locality of Daohugou (Inner Mongolia, China), Liminympha makarkini Ren & Engel and Daonymphes bisulca gen. et sp. n. This is the first record of this trait in the clade comprised of the superfamilies Myrmeleontoidea and Chrysopoidea. We interpret the recurrent veinlet in these species as a remnant of the condition present more basally in the psychopsoid + ithonoid + chrysopoid + myrmeleontoid clade (i.e., as a plesiomorphy). Other venational character states of D. bisulca of interest include the configuration of subcosta anterior (ScA), which is very similar to that of extant Nymphidae. We consider the short ScA terminating on ScP to be an autapomorphy of Neuroptera.

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