The lichen genus Henrica (Verrucariaceae, Eurotiomycetes) in northern Europe

In a previous three gene phylogeny study (nuITS rDNA, nuLSU rDNA and RPB1 region A-D) a well supported clade, most closely related to Verrucaria rupestris and also close to Sporodictyon and Atla, was found. Here this clade is identified as Henrica. A phylogeny of these genera based on the nuITS rDNA is supplied, and also a key to crustose species in Verrucariaceae with large, brown muriform spores. Based on material from northern Europe, Henrica is emended to include H. theleodes and H. melaspora, both new combinations in the previously monotypic genus. Verrucaria theleodes and V. scotinospora are lectotypified.