The oldest azhdarchoid pterosaur from the Late Jurassic Solnhofen Limestone (Early Tithonian) of Southern Germany

Based on an almost complete three-dimensionally preserved skeleton, a new genus and species of an azhdarchoid pterosaur Aurorazhdarcho primordius n.gen. n.sp. from the Late Jurassic Solnhofen limestone (Early Tithonian) of the Eichstatt area (Bavaria, Germany) is described. Furthermore, a new family the Protazhdarchidae is proposed. The specimen is attributed to the Azhdarchoidea based on its glenoid fossa level with the sternum, the shovel-like shape of the sternal plate, the wide furca of the coracoid, the metacarpus being longer than radius and ulna, the femur being 1/3 longer than the humerus, the femorotibial ratio, and the hammer-shaped humerus among other diagnostic features. Under UV-light, soft tissue preservation around the external mould of the head is visible. It consists of tiny flakes possibly remnants of skin. The dorsally curved outline of the external mould of the head suggests the presence of a cranial crest. The new species is the oldest record of the azhdarchoid pterosaurs. It supports the Eurasian origin of this group that includes the largest flying animal ever.