Nutritive value of winter-collected annual twigs of main European woody species, mistletoe and ivy and its possible consequences for winter foddering of livestock in prehistory

In Central Europe, forest grazing was probably the main way of providing winter feeding for livestock in prehistory; the main components of this diet most likely included annual twigs of woody species (Acer platanoides, Betula pendula, Carpinus betulus, Corylus avellana, Fagus sylvatica, Fraxinus excelsior, Picea abies, Pinus sylvestris, Populus tremula, Quercus robur, Salix caprea, Tilia cordata and Ulmus glabra), Hedera helix, Viscum album and senescent grassland biomass; however, their nutritive value has never been compared. Biomass samples were collected in the Czech Republic in February 2013 and analysed for concentration of nitrogen, phosphorus, potassium, calcium, magnesium, fibre and lignin. With the help of a recent analogy from the Altai Mountains (Russia), possible consequences of winter grazing on the development of forest vegetation and cattle breeds in prehistory were assessed. Hedera and Viscum were the best forage available in winter, and their nutritive value, according to the highest N and lowest fibre and lignin concentrations, was higher than the nutritive value of all winter-collected annual twigs of woody species. Nutritive value of annual twigs of all woody species was very low, especially compared with the quality of meadow hay, leaf-fodder or senescent steppe grassland biomass. Therefore, Hedera and Viscum might have been preferentially grazed by livestock and collected by ancient farmers for the supplementary feeding of privileged animals. According to recent analogy, annual twigs of woody species were browsed by livestock. Their insufficient quality could be one of the reasons for the low body size of cattle recorded in prehistory. The most realistic model of livestock breeding in prehistory seems to be year-round grazing, opening forests and causing deforestation in the surroundings of prehistoric settlements.